1. Department of Veterinary Anatomy, The University of Tokyo, Bunkyo-ku, Tokyo, Japan
2. Department of Experimental Animal Model for Human Disease, Center for Experimental Animals, Tokyo Medical and Dental University, Bunkyo-ku, Tokyo, Japan
3. Central Institute for Experimental Animals, Kawasaki-ku, Kawasaki, Kanagawa, Japan
4. Department of Neurobiology and Anatomy, The University of Utah, Salt Lake City, Utah, USA