1. Reproduction, Research Faculty of Agriculture, Hokkaido University, Sapporo, Hokkaido, Japan
2. Department of Endocrine Pharmacology, Tokyo University of Pharmacy and Life Sciences, Tokyo, Japan
3. School of Pharmaceutical Science, Fukushima, Japan
4. Global Station for Food, Land and Water Resources, Global Institution for Collaborative Research and Education, Hokkaido University, Sapporo, Hokkaido, Japan