1. Department of Stem Cells and Human Disease Models, Research Center for Animal Life Science, Shiga University of Medical Science, Otsu, Shiga, Japan
2. Institute for the Advanced Study of Human Biology (ASHBi), Kyoto University, Sakyo-ku, Kyoto, Japan
3. Department of Urology, Shiga University of Medical Science, Otsu, Shiga, Japan
4. Department of Evolutionary Studies of Biosystems, School of Advanced Sciences, SOKENDAI (The Graduate University for Advanced Studies), Hayama, Kanagawa, Japan
5. Department of Physiology, Shiga University of Medical Science, Otsu, Shiga, Japan
6. Department of Physiology, Keio University School of Medicine, Shinjuku-ku, Tokyo, Japan
7. PRESTO, Japan Science and Technology Agency, Honcho, Saitama, Japan