1. Yale School of Medicine Department of Genetics, , New Haven, CT , USA
2. Yale School of Medicine Department of Cellular and Molecular Physiology, , New Haven, CT , USA
3. Reproductive Biology Group, Division of Developmental Biology, Department of Biology, Faculty of Science, Utrecht University , Utrecht, Th e Netherlands
4. University of Amsterdam Center for Reproductive Medicine, Academic Medical Center, , Amsterdam, Th e Netherlands
5. Yale School of Medicine Department of Obstetrics, Gynecology and Reproductive Sciences, , New Haven, CT , USA
6. Yale School of Medicine Yale Cancer Center, , New Haven, CT , USA