1. Graduate School of Biostudies, Kyoto University, Kyoto, Japan
2. RIKEN Center for Sustainable Resource Science, Kanagawa, 230-0045 Japan
3. Graduate School of Pharmaceutical Science, Kyoto University, Kyoto, 606-8501 Japan
4. NODAI Genome Research Center, Tokyo University of Agriculture, Tokyo, 156-8502 Japan
5. Department of Bioscience, Tokyo University of Agriculture, Tokyo, 156-8502 Japan