1. Department of Forest Molecular Genetics and Biotechnology, Forestry and Forest Products Research Institute, Forest Research and Management Organization, 1 Matsunosato, Tsukuba, Ibaraki, 305-8687 Japan
2. School of Agriculture, Meiji University, 1-1-1 Higashi-Mita, Tama-ku, Kawasaki, Kanagawa, 214-8571 Japan
3. Forestry Research Institute, Hokkaido Research Organization, Koshunai, Bibai, Hokkaido, 079-0198 Japan
4. Faculty of Life and Environmental Sciences, University of Tsukuba, 1-1-1 Tennodai, Tsukuba, Ibaraki, 305-8572 Japan
5. Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, 1-1-1, Yayoi, Bunkyo-ku, Tokyo, 113-8657 Japan