1. Graduate School of Agricultural and Life Sciences, The University of Tokyo, 1-1-1 Yayoi, Bunkyo-ku, Tokyo, 113-8657 Japan
2. Hokkaido Agricultural Research Center, National Agriculture and Food Research Organization (NARO), 1 Hitsujigaoka, Toyohira-ku, Sapporo, Hokkaido, 062-8555 Japan
3. Division of Symbiotic Systems, National Institute for Basic Biology, 38 Nishigonaka, Myodaiji, Okazaki 444-8585, Aichi, Japan
4. Faculty of Agriculture, Shinshu University, Minamiminowa, Nagano, 399-4598 Japan
5. Graduate School of Life Sciences, Tohoku University, 2-1-1 Katahira, Aoba-ku, Sendai, 980-8577 Japan