1. Copenhagen Plant Science Center, Department of Plant & Environmental Sciences, University of Copenhagen , Frederiksberg C 1871 , Denmark
2. Max Planck Institute of Molecular Plant Physiology , Potsdam-Golm 14476 , Germany
3. Cereal Crops Research Unit, USDA-ARS , Madison, WI 53726 , USA
4. Department of Animal, Plant and Soil Sciences, School of Agriculture, Biomedicine and Environment, La Trobe University , Bundoora, VIC 3086 , Australia
5. Research School of Biology, The Australian National University , Acton, Australian Capital Territory 2600 , Australia
6. Joint International Research Laboratory of Metabolic & Developmental Sciences, State Key Laboratory of Hybrid Rice, SJTU-University of Adelaide Joint Centre for Agriculture and Health, School of Life Sciences and Biotechnology, Shanghai Jiao Tong University , Shanghai 200240 , China