1. State Key Laboratory of Crop Gene Exploration and Utilization in Southwest China, Sichuan Agricultural University , Chengdu 611130 , China
2. Rice Research Institute, Sichuan Agricultural University , Chengdu 611130 , China
3. State Key Laboratory of Southwestern Chinese Medicine Resources, and Innovative Institute of Chinese Medicine and Pharmacy, Chengdu University of Traditional Chinese Medicine , Chengdu 611137 , China