1. Key Laboratory of Aquatic Botany and Watershed Ecology, Wuhan Botanical Garden, Chinese Academy of Sciences , Wuhan 430074 , China
2. Center of Conservation Biology, Core Botanical Gardens, Chinese Academy of Sciences , Wuhan 430074 , China
3. College of Life Sciences, University of Chinese Academy of Sciences , Beijing 100049 , China
4. Wuhan Institute of Landscape Architecture , Wuhan 430081 , China
5. Department of Biology, Duke University , Durham, North Carolina 27708 , USA