1. School of Biological Sciences, Washington State University, Pullman, WA 99164-4236, USA
2. Biocenter University of Cologne, Institute for Plant Science, Cologne 50674, Germany
3. Max Planck Institute for Biology of Ageing, Cologne 50933, Germany
4. Max Planck Institute of Molecular Plant Physiology, Wissenschaftspark Golm, Potsdam 14476, Germany
5. Plant Biochemistry, Heinrich-Heine-University Düsseldorf, Düsseldorf D-40225, Germany