1. Horticultural Sciences, University of Florida , Gainesville, Florida 32611, USA
2. Institute of Plant Breeding, Genetics and Genomics, University of Georgia , Athens, Georgia 30602, USA
3. Department of Horticulture, University of Georgia , Athens, Georgia 30602, USA
4. Section of Plant Breeding and Genetics, Cornell University , Ithaca, New York 14853, USA
5. United States Department of Agriculture-Agricultural Research Service and Boyce Thompson Institute, Cornell University campus , Ithaca, New York 14853, USA