1. University of Chinese Academy of Sciences, Beijing 100049, China
2. National Key Laboratory for Plant Molecular Genetics, CAS Center for Excellence in Molecular Plant Sciences, Shanghai Institute of Plant Physiology and Ecology, Chinese Academy of Sciences, Shanghai 200031, China
3. Department of Crop Sciences, University of Illinois at Urbana Champaign, Urbana, Illinois 61801, USA
4. Department of Plant Biology, University of Illinois at Urbana Champaign, Urbana, Illinois 61801, USA