1. Tsukuba-Plant Innovation Center, University of Tsukuba , Tsukuba, Ibaraki 305-8572 , Japan
2. Department of Biochemistry, University of Oxford , Oxford, OX1 3RB , UK
3. Department of Biochemistry & Molecular Biology, Michigan State University , East Lansing, MI 48824 , USA
4. MSU-DOE Plant Research Laboratory, Michigan State University , East Lansing, MI 48824 , USA
5. Plant Resilience Institute, Michigan State University , East Lansing, MI 48824 , USA