1. Center for Sustainable Resource Science, RIKEN , 1-7-22 Suehiro-cho, Tsurumi, Yokohama, Kanagawa 230-0045 , Japan
2. Department of Life Sciences, The Milner Centre for Evolution, University of Bath , Bath BA2 7AY , UK
3. The Sainsbury Laboratory, Norwich Research Park , Norwich NR4 7UH , UK