1. The Key Laboratory of Biotechnology for Medicinal Plant of Jiangsu Province, School of Life Science, Jiangsu Normal University , Xuzhou, Jiangsu 221116 , China
2. Centre for Transformative Biotechnology of Medicinal and Food Plants, Jiangsu Normal University, Edinburgh University, Jiangsu Normal University , 101 Shanghai Road, Xuzhou 221116 , China
3. Institute of Molecular Plant Sciences, School of Biological Sciences, Edinburgh University , King's Buildings, Mayfield Road, Edinburgh EH9 3BF , UK