1. MOE Key Laboratory of Cell Activities and Stress Adaptations, School of Life Sciences, Lanzhou University , Lanzhou, Gansu 730000 , China
2. Institute of Plant Biology, Biological Research Centre, Eötvös Lóránd Research Network , Szeged , Hungary
3. Institute of Genetics and Biotechnology, Hungarian University of Agriculture and Life Sciences , Gödöllő , Hungary