1. Hokkaido Agricultural Research Center, National Agriculture and Food Research Organization (NARO), Sapporo, Japan
2. Division of Applied Genetics, Institute of Agrobiological Sciences, National Agriculture and Food Research Organization (NARO), Tsukuba, Japan
3. Research Faculty of Agriculture, Hokkaido University, Sapporo, Japan
4. Bioproduction Research Institute of Genome-based Biofactory, National Institute of Advanced Industrial Science and Technology (AIST), Sapporo, Japan