1. National Key Laboratory for Germplasm Innovation & Utilization of Horticultural Crops, College of Horticulture and Forestry Sciences, Huazhong Agricultural University , Wuhan 430070 , China
2. College of Horticulture, Northwest A&F University , Yangling 712100 , China
3. Hubei Hongshan Laboratory , Department of Science and Technology of Hubei Province, Wuhan 430070 , China