1. The Key Laboratory of Plant Development and Environmental Adaptation Biology, Ministry of Education, School of Life Sciences, Shandong University, Qingdao, 266237, China
2. Shandong Provincial Key Laboratory of Plant Stress, Shandong Normal University, Jinan, 250013, China
3. School of Life Sciences, Guangzhou University, Guangzhou, 510006, China
4. Noble Research Institute, LLC, Ardmore, Oklahoma, 73401
5. National Centre for Plant Gene Research (Beijing), Innovation Academy for Seed Design, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, 100101, China
6. College of Advanced Agricultural Sciences, University of Chinese Academy of Sciences, Beijing, 100049, China