1. Graduate School of International Agricultural Technology, Seoul National University , Pyeongchang 25354 , Korea
2. School of Biological Sciences and Technology, Chonnam National University , Gwangju 61186 , Korea
3. Crop Biotechnology Institute/GreenBio Science and Technology, Seoul National University , Pyeongchang 25354 , Korea
4. Department of Biology, De La Salle University , Manila 1004 , Philippines
5. LaSemilla Co. Ltd. , Pyeongchang 25354 , Korea