1. Bioscience and Biotechnology Center, Nagoya University , Nagoya 464-8601 , Japan
2. Graduate School of Bioagricultural Sciences, Nagoya University , Nagoya 464-8601 , Japan
3. School of Agriculture and Environment, The University of Western Australia , Crawley, WA 6009 , Australia
4. International Center for Research and Education in Agriculture, Nagoya University , Nagoya 464-8601 , Japan