1. Department of Horticulture, College of Horticulture and Landscape Architecture, Yangzhou University , Yangzhou, Jiangsu 225009 , China
2. State Key Laboratory for Managing Biotic and Chemical Threats to the Quality and Safety of Agro-products, Ministry of Agriculture and Rural Affairs Key Laboratory of Vegetable Legumes Germplasm Enhancement and Molecular Breeding in Southern China, Institute of Vegetables, Zhejiang Academy of Agricultural Sciences , Hangzhou 310021 , China
3. Department of Plant Science, School of Agriculture and Biology, Shanghai Jiao Tong University , Shanghai 200240 , China