1. CAS Key Laboratory of Tropical Plant Resources and Sustainable Use, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences , Kunming, Yunnan 650223 , China
2. State Key Laboratory for Conservation and Utilization of Bio-resources in Yunnan, Yunnan University , Kunming, Yunnan 650091 , China
3. University of Chinese Academy of Sciences , Beijing 100049 , China
4. Center of Economic Botany, Core Botanical Gardens, Chinese Academy of Sciences , Kunming, Yunnan 650223 , China