1. Molecular, Cellular & Integrative Physiology Program, University of California, Los Angeles, CA 90095-1606, USA
2. Department of Anatomy, Cell Biology & Physiology, Indiana University School of Medicine, Indianapolis, IN 46202, USA
3. Indiana Center for Musculoskeletal Health, Indiana University School of Medicine, Indianapolis, IN 46202, USA
4. Department of Integrative Biology and Physiology, University of California, Los Angeles, CA 90095-1606, USA
5. Department of Pathology and Laboratory Medicine, David Geffen School of Medicine at UCLA, University of California, Los Angeles, CA 90095, USA