1. Department of Medical Laboratory Science, College of Health Sciences, Debre Markos University , Debre Markos, Amhara Regional State , Ethiopia
2. Department of Microbiology, Immunology and Parasitology, College of Health Sciences, Addis Ababa University , Addis Ababa , Ethiopia
3. Armauer Hansen Research Institute , Addis Ababa , Ethiopia
4. Division of Experimental Medicine, University of California San Francisco , San Francisco, CA , USA
5. Department of Medicine, Division of Infectious Diseases and Emory Vaccine Center, Emory University School of Medicine, Emory University , Atlanta, GA , USA