1. Cologne Excellence Cluster on Cellular Stress Responses in Aging-Associated Diseases (CECAD), University of Cologne , Joseph-Stelzmann-Str. 26, 50931 Cologne , Germany
2. Institute of Genetics, University of Cologne , Zülpicher Str. 47a, 50674 Cologne , Germany
3. School of Biochemistry and Immunology, Trinity Biomedical Sciences Institute, Trinity College Dublin , 152-160 Pearse Street, Dublin, 2 , Ireland
4. Institute of Life Science, Swansea University , Swansea, SA2 8PP , UK