1. Department of Biology, Washington and Lee University , Lexington, VA , USA
2. Environmental Studies Program, Washington and Lee University , Lexington, VA, USA
3. Departmento de Ecología y Biología Evolutiva, Instituto de Investigaciones Biológicas Clemente Estable (IIBCE) , Montevideo , Uruguay
4. Departmento de Biodiversidad y Genética, Instituto de Investigaciones Biológicas Clemente Estable (IIBCE) , Montevideo , Uruguay
5. Environmental Studies Program, Washington and Lee University , Lexington, VA , USA
6. Department of Philosophy, Washington and Lee University , Lexington, VA , USA