1. Huazhong Agricultural University National Key Laboratory for Germplasm Innovation & Utilization of Horticultural Crops, , Wuhan 430070, China
2. Hubei Hongshan Laboratory, Wuhan 430070, China
3. Qinghai-Tibet Plateau Fruit Trees Scientific Observation Test Station (Ministry of Agriculture and Rural Affairs), Lhasa, Tibet 850032, China
4. Tibet Academy of Agricultural and Animal Husbandry Sciences Institute of Vegetables, , Lhasa, Tibet 850002, China