1. Hainan University Sanya Nanfan Research Institute, , Sanya, 572025, China
2. Yazhouwan National Laboratory , Sanya 572025, China
3. Huazhong Agricultural University National Key Laboratory for Germplasm Innovation & Utilization of Horticultural Crops, , Wuhan 430070, China
4. Huazhong Agricultural University National Key Laboratory of Crop Genetic Improvement and National Center of Plant Gene Research, , Wuhan 430070, China