1. Fujian Normal University Fujian Provincial Key Laboratory for Plant Eco-physiology, State Key Laboratory for Subtropical Mountain Ecology of the Ministry of Science and Technology and Fujian Province, College of Geographical Sciences, , Fuzhou 350007, China
2. Fujian Normal University College of Life Sciences, , Fuzhou 350117, China
3. Shunchang County Forestry Science and Technology Center of Fujian Province , Forestry Bureau of Shunchang, Shunchang 353200, China
4. Research Institute of Forestry, Fujian Research Institute of Forestry , Fuzhou 350000, China