1. Chinese Academy of Sciences Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, , Wuhan 430074, China
2. Chinese Academy of Sciences Center of Economic Botany, Core Botanical Gardens, , Wuhan 430074, China
3. University of Chinese Academy of Sciences , 19A Yuquanlu, Beijing, 100049, China
4. Wuhan University of Technology School of Chemistry, Chemical Engineering and Life Sciences, , Wuhan 430070, China