1. Department of Biomedical Informatics, Harvard Medical School , Boston , USA
2. Department of Statistics, University of Oxford , Oxford, UK
3. Hunan Provincial Key Lab on Bioinformatics, School of Computer Science and Engineering, Central South University , Changsha , China
4. Center for Medical Genetics, School of Life Sciences, Central South University , Changsha , China
5. School of Computer Science and Engineering, Nanyang Technological University , Singapore