1. Hadal & Nordcee, Department of Biology, University of Southern Denmark , Campusvej 55, Odense M 5230 , Denmark
2. Microbiology of Extreme Environments Laboratory, CNRS, IFREMER, Univ Brest , F-29280 Plouzané , France
3. Génomique Métabolique, Genoscope, Institut François Jacob, CEA, CNRS,University of Évry, Université Paris-Saclay , 91057 Evry , France
4. MARBEC, CNRS, IRD, Institut Français de Recherche pour L'Exploitation de la Mer, Univ Montpellier , 34200 Sète , France
5. Department of Ocean and Environmental Sciences , Tokyo University of Marine Science and Technology, 4-5-7 Konan, Minato-ku, Tokyo 108-8477 , Japan