1. Zhejiang University Laboratory of Systematic & Evolutionary Botany and Biodiversity, College of Life Sciences, , Hangzhou 310058, China
2. Henan University Laboratory of Plant Germplasm and Genetic Engineering, School of Life Sciences, , Kaifeng 475000, China
3. Zhejiang University Zhejiang Provincial Key Laboratory of Horticultural Plant Integrative Biology, , Hangzhou 31058, China
4. University of Wisconsin Department of Botany, , Madison, WI 53706, USA