1. Fudan University State Key Laboratory of Genetic Engineering and Fudan Center for Genetic Diversity and Designing Agriculture, School of Life Sciences, , Shanghai 200438, China
2. Fudan University Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, Institute of Biodiversity Science, , Shanghai 200438, China
3. Hunan Agricultural University College of Horticulture, , Changsha 410128, China