1. Institute of Biology and Soil Science, Far East Branch, Russian Academy of Sciences, Vladivostok 690022, Russia
2. Institute of Arctic Biology and Department of Biology and Wildlife, University of Alaska Fairbanks, Fairbanks, Alaska 99775, USA
3. Department of Biological Sciences, University of Alaska Anchorage, 3211 Providence Drive, Anchorage, Alaska 99508, USA
4. University of Alaska Museum, 907 Yukon Drive, University of Alaska Fairbanks, Fairbanks, Alaska 99775, USA
5. Department of Biology, Boston University, Boston, Massachusetts 02215, USA
6. Burke Museum of Natural History and Culture and Department of Biology, University of Washington, Seattle, Washington 98195, USA