1. Département des Sciences Fondamentales, Université du Québec à Chicoutimi, Canada
2. Department of Biology, Quebec Center for Biodiversity Science, McGill University, Montreal, Canada
3. Ministère de la Forêt, de la Faune et des Parcs du Québec, Direction de la recherche forestière, 2700 rue, Einstein, Canada
4. Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems, Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou, China