1. Centre of Excellence in Natural Resource Management, School of Agriculture and Environment, The University of Western Australia , 35 Stirling Terrace, Albany, WA 6330 , Australia
2. Biodiversity and Conservation Science, Department of Biodiversity, Conservation and Attractions , Locked Bag 104, Bentley Delivery Centre, Bentley, WA 6983 , Australia
3. School of Biological Sciences, The University of Western Australia , 35 Stirling Highway, Crawley, WA 6009 , Australia