1. Center for Ecological Research, Kyoto University , Otsu, Shiga 520-2113 , Japan
2. Department of Biological Sciences, Faculty of Agriculture and Life Science, Hirosaki University , 3 Bunkyo-cho, Hirosaki, Aomori 036-8561 , Japan
3. Forest Health Group, Kansai Research Center, Forestry and Forest Products Research Institute , 68 Nagaikyutaroh, Momoyama, Fushimi, Kyoto, Kyoto , Japan
4. Department of Forest Entomology, Forestry and Forest Products Research Institute , 1 Matsuno-sato, Tsukuba, Ibaraki 305-8687 , Japan