1. Department of Natural History Science, Graduate School of Science, Hokkaido University , Sapporo, Hokkaido 060 - 0810 , Japan
2. Department of Wildlife Biology, Forestry and Forest Products Research Institute , Tsukuba, Ibaraki 305-8687 , Japan
3. Department of Biology, School of Biological Sciences, Tokai University , Sapporo, Hokkaido 005 - 8601 , Japan
4. Department of Zoology, National Museum of Nature and Science , Tokyo 110-8718 , Japan