1. Department of Ecology & Evolutionary Biology, School of Biological Sciences, and Environment Institute, The University of Adelaide , Adelaide, South Australia 5005 , Australia
2. South Australian Museum , North Terrace, Adelaide, South Australia 5000 , Australia
3. Western Australian Museum , Welshpool DC, Western Australia 6986 , Australia
4. School of Biological Sciences, University of Western Australia , Crawley, Western Australia 6907 , Australia