1. State Key Laboratory of Integrated Management of Pest Insects and Rodents, Institute of Zoology, Chinese Academy of Science , Beijing 100101 , China
2. State Key Laboratory of North China Crop Improvement and Regulation, College of Plant Protection, Hebei Agricultural University , Baoding 071000 , China
3. Guangdong Key Laboratory of Animal Conservation and Resource Utilization and Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences , Guangzhou, Guangdong 510260 , China