1. Departamento de Genética, Ecologia e Evolução, Universidade Federal de Minas Gerais, Av. Antônio Carlos, 6627, Belo Horizonte, MG, 31270-901, Brazil
2. Natural Reserve System, University of California, Office of the President, Oakland,CA 94607-5200, USA
3. Centre of Excellence in Natural Resource Management, School of Agriculture & Environment, The University of Western Australia, 35 Stirling Terrace, Albany, WA 6330, Australia