1. Lewis E. Anderson Bryophyte Herbarium, Department of Biology, Duke University, Durham, NC 27708, USA
2. Department for BioMedical Research, University of Bern, Bern, Switzerland
3. Department of Biology, SUNY Oneonta, Oneonta, NY 13820, USA
4. Department of Biological Sciences, Texas Tech University, Lubbock, TX 79409, USA
5. Department of Natural History, NTNU University Museum, Norwegian University of Science and Technology, Trondheim, Norway