1. Palaeoscience Research Centre, School of Environmental and Rural Science, University of New England, Armidale,NSW 2351,Australia
2. Bavarian State Collection of Zoology, Bavarian Natural History Collections, Munich,Germany
3. Department Biology II, Ludwig-Maximilians-Universität München, Munich,Germany
4. GeoBio-Center, Ludwig-Maximilians-Universität München, Munich,Germany