1. Molecular Ecology and Conservation Genetics Laboratory, Department of Biological Applications and Technology, University of Ioannina , 45110 Ioannina , Greece
2. Ecological Genetics Research Unit, Organismal and Evolutionary Biology Research Programme, Faculty of Biological and Environmental Sciences, University of Helsinki , 00014 Finland
3. Area of Ecology and Biodiversity, The School of Biological Sciences, The University of Hong Kong , Hong Kong SAR