1. Wildlife Research Center , 3-3-7 Kotobashi, Sumida-ku, Tokyo 130-8606 , Japan
2. Laboratory of Plant Conservation Science, Faculty of Agriculture, Meijo University , Nagoya, Aichi 468-8502 , Japan
3. Faculty of Life and Environmental Sciences, University of Tsukuba , 1-1-1 Tennodai, Tsukuba, Ibaraki 305 - 8571 , Japan
4. Makino Herbarium, Tokyo Metropolitan University , 1-1 Minamiosawa, Hachioji, Tokyo 192 - 0364 , Japan