1. Biological Laboratory, Faculty of Education, Nagasaki University , Bunkyo, Nagasaki , Japan
2. Graduate School of Global Environmental Studies, Kyoto University , Yoshida Nihonmatsu-cho, Sakyo, Kyoto , Japan
3. Faculty of Human Environmental Studies, Hiroshima Shudo University , Hiroshima , Japan
4. Division of Environmental Science and Ecological Engineering, College of Life Sciences and Biotechnology, Korea University , Seoul 02841 , Republic of Korea
5. Department of Biology, Faculty of Science, Shinshu University , Matsumoto, Nagano , Japan