1. CNRS, Université Paul Sabatier, IRD, UMR 5174 EDB (Laboratoire Évolution & Diversité Biologique), Toulouse, France
2. Department of Animal and Plant Sciences, University of Sheffield, Western Bank, Sheffield, UK
3. Claude E. Phillips Herbarium, Delaware State University, Dover, DE, USA
4. Herbier, Département Flore, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar
5. Kew Madagascar Conservation Centre, Ivandry, Antananarivo, Madagascar
6. Comparative Plant and Fungal Biology, Royal Botanic Gardens Kew, Richmond, Surrey, UK